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ACSL4 was first identified as a navigator of ferroptosis by haploid genetic screening [13]

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Additionally, one study discovered the first ferroptosis defense mechanism operating within mitochondria: dihydroorotate dehydrogenase (DHODH), an enzyme involved in pyrimidine synthesis, converts CoQ to CoQH2 in the inner mitochondrial membrane, neutralizing the lipid peroxidation defense against ferroptosis in mitochondria (Mao et al., 2021)